15 de septiembre de 2009

Aspectos Generales del Phylum Placozoa

Los placozoos son de los organismos más simples descritos a la fecha. En realidad, se conoce poco acerca de ellos, y no han sido observados en su hábitat natural. Nadie sabe en cuál tipo de sustratos se encuentran, de que se alimentan, como se reproducen, cuantas especies hay en realidad, en fin son un grupo desconocido.

Estos organismos fueron descubiertos en 1883, por zoólogo Alemán Franz Eilhard Schulze, el cual los observó viviendo en un acuario. A partir de ahí, lo que se ha podido investigar acerca de ellos, ha sido, unas pocas observaciones que se han realizado en colonias que se han mantenido vivas bajo condiciones de laboratorio, o muy pocos casos en su hábitat natural.

Distribución

El grupo esta distribuido en la región tropical y subtropical.  En el Mar mediterráneo, Mare Rojo, Océano Caribe, Golfo de México, Samoa Oriental, Japón, Hawaii y la Gran Barrera de Arrecifes. Los zoólogos creyeron, al inicio, que se trataba de la larva de alguna especie de medusa aún no descrita, pero investigaciones posteriores demostraron que es un organismo adulto, posiblemente el metazoario viviente más primitivo.

Distribución Geográfica de Trichoplax adhaerens 


Morfología

Los placozoos o placozoarios son microorganismo pluricelulares y en forma de ameba ( ameboide ), miden entre 1 - 2 mm de diámetro; transparentes con tonos rosa claros, su cuerpo es aplanado, su estructura corporal es muy sencilla. Están formado por dos capas epiteliales (dorsal y ventral) separadas por otra capa simple de células sueltas capaces de contraerse ( algo similar a un músculo ). Poseen dos tipos de células; cilíndricas ciliadas y glandulares no ciliadas que constituyen la porción ventral del cuerpo. La porción dorsal la constituyen únicamente células epiteliales, las cuales son ciliadas y aplanadas y que recubren su parte externa. No hay simetría corporal ni órganos diferenciados; carece, asimismo, de sistemas nervioso y muscular.



Animación de Trichoplax adhaerens, fuente: Tree of Life Project

Fisiología

Su alimentación se lleva a cabo por medio de absorción, a través de su superficie ventral, por medio de predigestión extracelular y se presume que se alimenta de algas y protozoos.

Clasificación

La clasificación biológica está poco desarrollada, solamente se reconoce una familia, Trichoplacidae, y se reconocen dos géneros con una sola especie cada uno: Trichoplax adhaerens y Treptoplax reptans



Treptoplax reptans, se describió en 1896 y no ha vuelto a verse desde entonces.   Aunque muy poco es conocido de los Placozoos, investigaciones usando variaciones genéticas y marcadores moleculares han concluido que pudiera existir una variedad mas amplia de especies dentro de este filo.

Reproducción y Desarrollo

Trichoplax se reproduce de dos maneras: por fisión binaria y por gemación. Sin embargo se ha reportado la producción de óvulos y espermios, Sin embargo no se observan órganos sexuales.

Trichoplax adhaerens

Su desarrollo se ha observado únicamente hasta el estado de 64 células, donde se llega al punto en que las células cesan sus divisiones mitóticas, pero el ADN continúa multiplicándose hasta que los núcleos se abren.

Relaciones Evolutivas

Algunos científicos presumían que los placozoos podrían haber sido el ancestro común de muchos phyla primitivos, por su gran simplicidad, ya que es sin duda el metazoo conocido más simple, además el contenido de ADN en el núcleo de sus células es comparativamente menor que otros animales.

Sin embargo, el tipo de uniones celulares de sus células epiteliales ( por proteínas extra celulares ), la cual es una condición presente en todos los animales, exceptuando las esponjas, sugirió que probablemente no sean tan primitivos.

Información obtenida a partir del análisis de secuencias de ADN ( 18S), han indicado que los placozoos evolucionaron aún más tarde en la historia. Esto incluiría que los placozoos al evolucionar, simplificaron su sistema nervioso.


Cladograma que explica la posición taxonómica de Placozoa, tomado de Schierwater et al. 2009
 
Para leer más

Schierwater et al. 2009. Concatenated Analysis Sheds Light on Early Metazoan Evolution and Fuels a Modern ‘‘Urmetazoon’’ Hypothesis. PLoS Biology January 2009 Volume 7, Issue 1. Disponible en:
http://www.ecolevol.de/index.php?option=com_content&task=view&id=249&Itemid=102


Dellaporta et al. 2002. A Case for Sequencing the Trichoplax Genome. Disponible en:
http://www.genome.gov/Pages/Research/Sequencing/SeqProposals/TrichoplaxSEQ021203.pdf
 
Schierwater B, 2005. My favorite animal, Trichoplax adhaerens. BioEssays 27:1294-1302. Disponible en :
http://pharyngula.org/index/weblog/comments/mysterious_trichoplax/ 
A NEW RECORD OF Trichoplax adhaerens F. E. SCHULZE (PHYLUM PLACOZOA) IN THE MEXICAN CARIBBEAN SEA SCIENTIFIC NOTE

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"Su desarrollo se ha observado únicamente hasta el estado de 64 células, donde se llega al punto en que las células cesan sus divisiones mitóticas, pero el ADN continúa multiplicándose hasta que los núcleos se abren."

No entiendo bien ese párrafo, ¿quiere decir, que una vez terminadas las mitosis las celulas quedan en una especie de fase S?

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Review this sports season to be hit into 4 powerful force of thing ministry final, thunderbolt has spent big price to eat the Jin Si that enter handkerchief, the ox also has the doughty planted agent of Nuo A, Buzeer. Only lake person and showing tremendous enthusiasm are next Yao Ming's potential homes. Think carefully, miami is optimal choice of Yao Ming undoubtedly, have " 3 tycoons " showing tremendous enthusiasm, the urgentest need fills strong is planted agent, er of big Z, the root bark of the tree peony year the work is expensive already and injury is ceaseless, if Yao Ming can sacrifice,pay joins in showing tremendous enthusiasm, comprise bloodcurdling " 4 tycoons " , so he is very possible dream of champion of one round NBA. Besides, lake person also will be the 2nd choice of Yao Ming, above all los angeles is to be able to be rivalled cease Si Dui's city, once Yao Ming is entered advocate " the city of angel " , have tremendous stimulative effect undoubtedly in trade development to its, be in early 2005 when, division comparing ever invited Yao Ming one case personally " achieve great undertaking in all " . If once Yao Ming joins in, he will obtain the space of very good play, mix in competition ground trade hopeful obtain " double bumper harvest " .

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Sikeersi today Hei Bei of 37 2 balls searchs Xia Huoyong Bie Manlian painstakingly shoot boots
Sikeersi today Hei Bei of 37 2 balls searchs Xia Huoyong Bie Manlian painstakingly shoot boots
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Graceful coupletIn field veteranSikeersiAfter knee got hurt January, fail to reappear all the time, his contract today summer expire, graceful wing grows Brian before - Luobusen thinks Sikeersi is probable at this point hang up one's spikes, reappear no longer. Young generalWeierbeikeThis sports season carried extremely high attendance rate, but he of department office forward advances 2 balls 37 times only, hei Bei thinks this is the cause that he often gues-stars frontier way.
After announcing to retire Spring 2011, the unexpectedly announced Sikeersi in the winter last year reappear, decide to kick a year again. However this sports season his condition how terrible, andJi GesiThe field in graceful couplet is clear when coming on the stage at the same time lose power and influence, then Ferguson emphasizes education young generalKelaiweili, and Sikeersi disappears for long already in fan line of sight, he comes on the stage last time still was on January 16Sufficient total cupBe a guestMu of the Western Han Dynastythe match, also get hurt accordingly.
Bisikeersi is big Jigesi of a year old already decided to renew the contract one year, the coming year again battle, [url=http://wholesalenikeshoes.manifo.com/]wholesale nike shoes[/url] the ginger head summer that Luo Busen of header of blood of iron of Dan Qianman couplet worries about 38 years old is possible forever hang up one's spikes: "Ji Gesi will kick a year again, he looks and when 21 years old same health. I am not affirmatory to Sikeersi, his knee appeared a few problems, so I think he retires likely. Only he knows to whether can fall to reappear again in emergency, but his manner and training are very active, so he still had better stay. So he still had better stay..
Differ with Sikeersi, shellfish of Er of young general dimension overcomes this sports season attendance rate is extremely high, and suffer Ferguson to trust quite, but 37 his matches just advance 2 balls, goal of this group of sports season nation still compares the goal in graceful couplet much, face an one foot to suffer intense doubt. Hei Bei advances 1 ball only in league matches this year, that or last year in October advocate field is rightSituoke cityWhen head burst open the door, additionally he still is inOu GuanBe a guestEmperor horseWhen head burst open the door, win wide recognition.
Because want to gues-star wing,Hei Bei professes not to score a goal is: "I miss more ball, my performance is more steady. I did not kick the niche that I like all the time, play different position however, this makes me right other charge the position has closer understanding, and this can let you be benefited only. " the graceful couplet that graceful city heart of Zhou Yi is compared is Hei Bei 100 milepost, he spoke of the Europe coronal goal to emperor horse: "This is a great day to club and me. Episcopal practice chose a lineup that can take fraction in Bainawu, and we have an opportunity to win victory. We go out regretfully finally bureau, but I am in to oneself Bainawu's performance is very satisfactory, I won't forget this experience forever. "

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